Fission–fusion Society
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ethology Ethology is the scientific study of animal behaviour, usually with a focus on behaviour under natural conditions, and viewing behaviour as an evolutionarily adaptive trait. Behaviourism as a term also describes the scientific and objectiv ...
, a fission–fusion society is one in which the size and composition of the social group change as time passes and animals move throughout the environment; animals merge into a group (fusion)—e.g. sleeping in one place—or split (fission)—e.g. foraging in small groups during the day. For species that live in fission–fusion societies, group composition is a dynamic property. The change in composition, subgroup size, and dispersion of different groups are 3 main elements of a fission-fusion society. This
social organization In sociology, a social organization is a pattern of relationships between and among individuals and social groups. Characteristics of social organization can include qualities such as sexual composition, spatiotemporal cohesion, leadership, s ...
is found in several primates,
elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae an ...
s,
cetacea Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel them ...
ns,
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
s, social
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
s, some birds and some fish.


Species

Fission-fusion societies occur among many different species of primates (e.g.
chimpanzees The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative th ...
,
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genu ...
s, and
human Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
s), elephants (e.g.
forest elephants The African forest elephant (''Loxodonta cyclotis'') is one of the two living African elephant species. It is native to humid forests in West Africa and the Congo Basin. It is the smallest of the three living elephant species, reaching a should ...
, African elephants), and
bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most ...
s (e.g. northern long-eared bats). The change in composition, subgroup size, and dispersion of different groups are 3 main elements of a fission-fusion society.


Primates


Chimpanzees

Chimpanzees often form smaller subgroups when travelling for longer periods at a time in between each food patch. When obtaining food, the size of subgroups can change depending on how much food is available and how far away the food may be. If food is worth retrieving due to little travel costs, subgroup size will enlarge. So among chimpanzees, the abundance of food and how dense it may be are factors that contribute to the changes of subgroup sizes.


Orangutans

Orangutans are one type of primates that model individual-based fission-fusion. Travel parties are established among this species inhabiting specifically in a Sumatran forest, and there are several benefits. Mating opportunities are a large benefit of grouping, as parties are most substantial during high mating activity. Infant socialization also contains benefits as well as costs, due to their needs to be cared for. Females are required to carry their infants, and those with infants of mid-size experience greater costs than those of small sizes. Carrying a small infant does not require much, and they become less dependent as they begin to wean. Mid-sized infants on the other hand, require the most energy. When travelling, females are required to support their mid-sized infants by carrying them, and waiting for them if they've fallen behind.


Humans

Humans also form fission-fusion societies. In hunter-gatherer societies, humans form groups which are made up of several individuals that may split up to obtain different resources. Another example of a fission-fusion society in hunter-gatherer societies is communication among the group. Groups may begin to split due to arguments and disagreements. Among humans, gossip and language in general is also an important feature involved in fission-fusion. Communication keeps distant groups together although they may not be within close distances of each other.


Elephants

Elephants display grouping as it helps to minimize risks and enhance benefits. Forest elephants often fuse into larger groups throughout forest clearings, to exchange information and maximize social opportunities. Elephants are drawn to large parties present at forest clearings, and remain in the clearing for a longer period of time if there are individuals outside of their party present. Young African male elephants display a preference for larger groups, in order to communicate with other elephants and to explore dominance. Adolescent males can gain knowledge from adult males and acquire information about their new social methods.


Bats

Bats are one of the orders of animals which shows an advanced fission-fusion society. Among female northern long-eared bats, switching roosts is common. There are several factors involved when switching roosts, which can include canopy cover and height, decay stage of the roost, and tree height. Geographic regions contribute to the switching of roosts, as females have been shown to switch when temperatures rise in Kentucky, and less when in a colder climate in Nova Scotia. There are also three important behaviours involved in roost-switching, which are fission-fusion grouping, synchronized movement, and settlement behaviour. Settlement behaviour is when bats remain in the most desirable roost possible, synchronized movement is when bats choose to move to another roost in a synchronized manner, and fission-fusion behaviour is when a bat colony separates into sub-colonies which then combine back together to form a large colony. A reason that bats can display fission-fusion behaviours is due to the risk of infection. Increased risk of disease can occur from settlement and synchronized behaviours, but fission-fusion societies are capable of reducing the risk of disease. Fission-fusion societies are capable of reducing the risk because of the frequent separation into subgroups.


Structure

These societies change frequently in their size and composition, making up a permanent social group called the "parent group". Permanent social networks consist of all individual members of a faunal community and often varies to track changes in their environment and based on individual animal dynamics. In a fission–fusion society, the main parent group can fracture (fission) into smaller stable subgroups or individuals to adapt to
environmental A biophysical environment is a biotic and abiotic surrounding of an organism or population, and consequently includes the factors that have an influence in their survival, development, and evolution. A biophysical environment can vary in scale f ...
or social circumstances. For example, a number of males may break off from the main group in order to hunt or forage for food during the day, but at night they may return to join (fusion) the primary group to share food and partake in other activities. Overlapping of so-called "parent groups" territorially is also frequent, resulting in more interaction and mingling of community members, further altering the make-up of the parent group. This results in instances where, say, a female chimpanzee may generally belong to one parent group, but encounters a male who belongs to a neighboring community. If they
copulate Sexual intercourse (or coitus or copulation) is a sexual activity typically involving the insertion and thrusting of the penis into the vagina for sexual pleasure or reproduction.Sexual intercourse most commonly means penile–vaginal penetr ...
, the female may stay with the male for several days and come into contact with his parent group, temporarily "fusing" into the male's community. In some cases, animals may leave one parent group to associate themselves with another, usually for reproductive reasons.


See also

*
Pair bonding Pair or PAIR or Pairing may refer to: Government and politics * Pair (parliamentary convention), matching of members unable to attend, so as not to change the voting margin * ''Pair'', a member of the Prussian House of Lords * ''Pair'', the Frenc ...
*
Human bonding Human bonding is the process of development of a close, interpersonal relationship between two or more people. It most commonly takes place between family members or friends, but can also develop among groups, such as sporting teams and whenever ...


References

{{DEFAULTSORT:Fission-fusion society Group processes Zoosemiotics Ethology